Flik as a hero was insanely smart and took his invention to incredible levels. Once Hopper and his clan attacked, Flik was able to not only hold them off, but fully and successfully defend his colony as their hero.
The villainous Flik was more of a biological inventor. Always creating potions and elixirs, trying to become stronger. He was eventually successful and became to truest form of a fire ant. Letting the power go to his head, Flik killed off Hopper, his crew, and took the entire ant colony with it. Now Flik wanders alone incinerating and bug in his path.